Phyllostegia warshaueri

H.St.John (1987)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Lamiaceae Genus: Phyllostegia

Description

Key Characters:

Growth Form: Scandent to decumbent lianas.

Stems: Stems ca. 10–30 dm long, antrorsely puberulent.

Roots:

Leaves: Leaves simple. Opposite. Blades oblong–elliptic to ovate or oblong–ovate, 7.5–15 cm long, 2–6.6 cm wide. Apex acute. Base cuneate. Surfaces glabrate except lower surfaces occasionally sparsely puberulent on margins or veins. Margins serrate, occasionally remotely so. Petioles 2–5 cm long, glabrous or with a few scattered hairs. Stipules absent.

Flowers: Flowers 6–14 per verticillaster, in unbranched, elongate, terminal, racemose inflorescences up to 20 cm long, pedicels 25–40 mm long, glabrous or with a few hairs at base, bracts ovate to elliptic, 7–21 mm long, usually sparsely ciliate, lowermost pair larger, 11–50 mm long Flowers 1–14 or rarely more per verticillaster, on short or elongate pedicels, rarely these on a common peduncle, arranged in terminal, bracteate, racemose inflorescences on the main and lateral stems, either simple and unbranched or compound, formed by the principal inflorescence flanked by smaller secondary or even tertiary, lateral, racemose inflorescences. Calyx narrowly obconical, ca. 6–8 mm long at anthesis, enlarging in fruit, nerves inconspicuous, glabrous, usually 5-toothed, the teeth deltate, unequally divided, ca. 2–3 mm long, apex rounded to obtuse, usually ciliate, splitting longitudinally, usually persistent or sometimes only ventral 1/2 persistent. Corolla bilabiate, white, upper lip dark rose, tube narrowly funnelform, 18–20 mm long, sparsely villous on dorsal surface, upper lip ca. 6 mm long entire or 2-lobed, ciliate, lower lip ca. 12–15 mm long, 3-lobed, lateral lobes ovate, slightly asymmetrical, lower lobe broadly ovate to suborbicular, slightly folded to form a groove, margins somewhat undulate, apex emarginate or entire; nectary disk often present at base of ovary, annular or developed on anterior side only. Stamens 4, both pairs of similar length or upper pair slightly longer, ascending the upper corolla lip, included, aligned with the sinuses; filaments inserted near throat, usually pubescent; anthers dithecal, opening by longitudinal slits; anther sacs divaricate. Ovary superior, 2-carpellate, placentation basal-axile; ovules 1 per ovary lobe; style 1, included, arising between lobes, shortly 2-lobed, the lobes clavate; stigmatic only at apex, 1 lobe often rudimentary; stigmas 2.

Fruit: Nutlets ca. 6–7 mm long. Seeds with endosperm absent or scanty and oily.

Ploidy:

Habitat: Known from scattered collections in wet forest.

Elevation Range: 720–950 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Phytologia 63: 182 (1987)

Other References

Wagner et al. 1990:826 (H); Wagner 1999a:277

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Metrosideros- Cheirodendron Montane Wet Forest. PRESERVED_SPECIMEN K. R. Wood 2190 Hawaii PTBG 12/18/1992
2 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Lydgate, J.M. s.n. Maui BISH
3 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN S. P. Perlman & L. Perry 21353 Hawaii US 1/20/2009
4 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Perlman, S. P.; Perry, L. 21353 Hawaii US 1/20/2009
5 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wood, K. R.; Perlman, S. P.; Leialoha, J.; Stevens, B.; Jokiel, G. 4119 Hawaii US 3/7/1995
6 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN s. p. perlman, k. wood & r. warshauer 14691 Hawaii US 3/6/1995
7 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN k. r. wood & s. p. perlman 2190 Hawaii US 12/18/1992
8 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Perlman, S. P.; Wood, K.; Warshauer, R. 14691 Hawaii US 3/6/1995
9 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN k. r. wood, s. p. perlman, j. leialoha, b. stevens & g. jokiel 4119 Hawaii US 3/7/1995
10 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN s. p. perlman, k. wood & b. stevens 14185 Hawaii US 5/12/1994
11 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wood, K. R.; Perlman, S. P. 2190 Hawaii US 12/18/1992
12 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN s. p. perlman & l. perry 21353 Hawaii US 1/20/2009
13 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Perlman, S. P.; Wood, K.; Stevens, B. 14185 Hawaii US 5/12/1994
14 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information In forest near Cyanea platyphylla with Acacia koa-Metrosideros forest with Psychotria, Cyanea, Clermontia, Cheirodendron, Melicope, Coprosma, Clidemia hirta, Pipturus, Freycinetia. PRESERVED_SPECIMEN perlman, s.p. wood, k.r.; stevens, b. collector number: 14185 Hawaii BISH 5/12/1994
15 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Corn, C. ESP 379 Hawaii BISH 2/11/1982
16 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Manoa Cliffs PRESERVED_SPECIMEN anderson, s.j. collector number: 470 Oahu BISH 2/11/1982
17 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN anderson, s.j. collector number: 470 Molokai BISH 2/11/1982
18 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Growing with in 50 ft. of Blair road. Habitat: open Acacia koa - Metrosideros forest with understory of Cibotium, Cheirodendron, Coprosma, Antidesma, Psychotria, Gouldia, ect. Groundcover dominated by Athyrium sandwichianum. PRESERVED_SPECIMEN Davis, J. Collector Number: 602 Hawaii BISH 9/8/1980
19 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Climbing on Athyrium sandwichianum, Cibotium, Metrosideros, Antidesma, and Coprosma in the open `ohi`a west of Kilau Stream. PRESERVED_SPECIMEN Anderson, S.J. Collector Number: 470 Hawaii BISH 2/11/1982
20 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Area 50 X 75 ft, patch of vines climbing to 20' on Athyrium sandwichianum, Cibotium, Metrosideros, Antidesma, and Coprosma in open ohia forest. PRESERVED_SPECIMEN Anderson, S.J. 470 Hawaii BISH 2/11/1982
21 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN anderson, s.j. clarke, g.; cuddihy, l.w.; davis, j. collector number: 421 Molokai BISH 3/3/1981
22 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN lydgate, j.m. collector number: s.n. Maui BISH
23 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Growing with in 50 ft. of Blair road. Habitat: open Acacia koa - Metrosideros forest with understory of Cibotium, Cheirodendron, Coprosma, Antidesma, Psychotria, Gouldia, ect. Groundcover dominated by Athyrium sandwichianum. PRESERVED_SPECIMEN Davis, J. 602 Hawaii BISH 9/8/1980
24 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN corn, c. collector number: esp 379 Hawaii BISH 2/11/1982
25 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Open Acacia - Metrosideros forest. PRESERVED_SPECIMEN anderson, s.j. clarke, g.; cuddihy, l.w.; davis, j. collector number: 421 Hawaii BISH 3/3/1981
26 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Trailing across a small stream. PRESERVED_SPECIMEN rock, j.f.c. collector number: 8380 Hawaii BISH 6/1/1910
27 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Open Acacia - Metrosideros forest. PRESERVED_SPECIMEN Anderson, S.J. Clarke, G.; Cuddihy, L.W.; Davis, J. Collector Number: 421 Hawaii BISH 3/3/1981
28 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Open Acacia - Metrosideros forest. PRESERVED_SPECIMEN Anderson, S.J. 421 Hawaii BISH 3/3/1981
29 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Tomich, Michael FRW 4005 Hawaii BISH 5/28/1979
30 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Tanich, M. FRW 4005 Hawaii BISH 5/28/1979
31 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information In Acacia koa - Metrosideros mesic forest with Pipturus albidus, Cibotium, Perrottetia sandwicensis, Rubus hawaiensis, Clermontia parviflora, Broussaisia arguta, Urera glabra, Antidesma platyphylla, Freycinetia arborea, Cheirodendron trigynum, Ilex anomala, Astelia menziesiana, Peperomia. PRESERVED_SPECIMEN Perlman, S.P. et al. Collector Number: 21353 Hawaii BISH 1/20/2009
32 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Open Acacia - Metrosideros forest. PRESERVED_SPECIMEN Anderson, S.J. Clarke, G.; Cuddihy, L.W.; Davis, J. Collector Number: 421 Hawaii BISH 3/3/1981
33 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Growing with in 50 ft. of Blair road. Habitat: open Acacia koa - Metrosideros forest with understory of Cibotium, Cheirodendron, Coprosma, Antidesma, Psychotria, Gouldia, ect. Groundcover dominated by Athyrium sandwichianum. PRESERVED_SPECIMEN davis, j. collector number: 602 Hawaii BISH 9/8/1980
34 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information In Acacia koa - Metrosideros mesic forest with Pipturus albidus, Cibotium, Perrottetia sandwicensis, Rubus hawaiensis, Clermontia parviflora, Broussaisia arguta, Urera glabra, Antidesma platyphylla, Freycinetia arborea, Cheirodendron trigynum, Ilex anomala, Astelia menziesiana, Peperomia. PRESERVED_SPECIMEN perlman, s.p. et al. collector number: 21353 Hawaii BISH 1/20/2009
35 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information In Acacia koa - Metrosideros mesic forest with Pipturus albidus, Cibotium, Perrottetia sandwicensis, Rubus hawaiensis, Clermontia parviflora, Broussaisia arguta, Urera glabra, Antidesma platyphylla, Freycinetia arborea, Cheirodendron trigynum, Ilex anomala, Astelia menziesiana, Peperomia. PRESERVED_SPECIMEN Perlman, S. 21353 Hawaii BISH 1/20/2009
36 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Growing on the steep southern bank of unnamed stream just south of Humuula Trail. Growing with METPOL, ACAKOA, ILEANO, CHETRI, CCOPspp., CIBspp., PSICAT. PRESERVED_SPECIMEN Loo, R. HA-OOK-A-3337 Hawaii BISH 6/20/2017
37 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN warshauer, f.r. (sub warshauer) collector number: frw 4005 Hawaii BISH 5/28/1979
38 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Open Acacia - Metrosideros forest. PRESERVED_SPECIMEN anderson, s.j. clarke, g.; cuddihy, l.w.; davis, j. collector number: 421 Hawaii BISH 3/3/1981
39 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Trailing across a small stream. PRESERVED_SPECIMEN Rock, J.F.C. 8380 Hawaii BISH 6/1/1910
40 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Open Acacia-Metrosideros forest, vining to 12', 10'x20' area PRESERVED_SPECIMEN anderson, s.j. clarke, g.; cuddihy, l.w.; davis, j. collector number: 421 Hawaii BISH 3/3/1981
41 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Open Acacia-Metrosideros forest, vining to 12', 10'x20' area PRESERVED_SPECIMEN Anderson, S.J. 421 Hawaii BISH 3/3/1981
42 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron montane wet forest w/ Sadleria squarrosa, Peperomia macraeana, Dubautia plantaginea, Melicope clusiifolia, Broussaisia arguta, Cyrtandra platyphylla. PRESERVED_SPECIMEN wood, k.r. perlman, s.; leialoha, j.; stevens, b.; jokiel, g. collector number: 4119 Hawaii BISH 3/7/1995
43 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron montane wet forest; steep forests of ferns and trees, overlooking deep valley. PRESERVED_SPECIMEN wood, k.r. perlman, s. collector number: 2190 Hawaii BISH 12/18/1992
44 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Open Acacia - Metrosideros forest. PRESERVED_SPECIMEN Anderson, S.J. 421 Hawaii BISH 3/3/1981
45 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information In forest near Cyanea platyphylla with Acacia koa-Metrosideros forest with Psychotria, Cyanea, Clermontia, Cheirodendron, Melicope, Coprosma, Clidemia hirta, Pipturus, Freycinetia. PRESERVED_SPECIMEN Perlman, S. 14185 Hawaii BISH 5/12/1994
46 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Climbing on Athyrium sandwichianum, Cibotium, Metrosideros, Antidesma, and Coprosma in the open `ohi`a west of Kilau Stream. PRESERVED_SPECIMEN Anderson, S.J. 470 Hawaii BISH 2/11/1982
47 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron montane wet forest; steep forests of ferns and trees, overlooking deep valley. PRESERVED_SPECIMEN Wood, K.R. 2190 Hawaii BISH 12/18/1992
48 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information In Metrosideros-Acacia koa lowland mesic forest. PRESERVED_SPECIMEN Steve Perlman 14691 Hawaii PTBG 3/6/1995
49 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information In forest near Cyanea platyphylla. PRESERVED_SPECIMEN Steve Perlman 14185 Hawaii PTBG 5/12/1994
50 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Acacia koa- Metrosideros- Dicranopteris linearis wet forest. PRESERVED_SPECIMEN Steve Perlman 23091 Hawaii PTBG 11/7/2012
51 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron Montane Wet Forest. PRESERVED_SPECIMEN K. R. Wood 4119 Hawaii PTBG 3/7/1995
52 Phyllostegia warshaueri H.St.John Contact Bishop Museum Botany department for locality information Acacia koa- Metrosideros mesic forest. PRESERVED_SPECIMEN Steve Perlman 21353 Hawaii PTBG 1/20/2009